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Here is the cell style data for my custom version of Section:

 CellFrame->{{0, 0}, {0, 0}},
 CellMargins->{{10, Inherited}, {8, Inherited}},

But when I type command+1 I still get the old red Title cell.

enter image description here

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up vote 7 down vote accepted

You'll also need to make a version of the Title cell which has the MenuCommandKey option unset. I.e.,

Cell[StyleData["Title"], MenuCommandKey->None]

The FE doesn't have a standard way of resolving styles with overlapping MenuCommandKey options, so if you want it to work properly, it's important to make sure that you prevent overlaps.

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